Department of Fisheries
and Aquatic Sciences

 
Karl E. Havens

Professor and Chair

B.A. Biology, State University of New York at Buffalo, 1979

M.S. Biology, West Virginia University, 1981

Ph.D. Biology, West Virginia University, 1984

 Karl Havens is Chair of the Department of Fisheries and Aquatic Sciences, where he provides program leadership for research, education, and extension activities that support the missions of the Department, IFAS, and the University. He is an applied limnologist whose research focus is on trophic interactions in lake plankton communities, interactions between aquatic plants, plankton and fish, and the ecology and management of shallow eutrophic lakes. He has conducted collaborative research projects with scientists in Brazil, Canada, China, Denmark, Italy, Japan and the Netherlands, and is actively involved in the application of science in the management of Florida lakes.

Prior to joining the Department in December 2004, Dr. Havens was Chief Environmental Scientist at the South Florida Water Management District, where he led a rigorous limnological research program, worked with lake managers to apply research results in large-scale restoration projects, and communicated with decision makers, elected officials, representatives of other state and federal agencies, and the general public. Earlier in his career, he was a faculty member in the Department of Biological Sciences and Water Resources Research Institute at Kent State University where he taught a variety of courses in biology, ecology, and aquatic sciences, carried out research dealing with plankton dynamics in temperate lakes, and mentored graduate students in plankton ecology and limnology.

In 1999, Dr. Havens was presented the Edward Deevey, Jr. Award from the Florida Lake Management Society for his "contribution to the scientific knowledge of Lake Okeechobee." He is active in professional service as Associate Editor of Environmental Pollution, Scientific Advisory Board Member for the Korean Journal of Limnology, Chapter President for Sigma Xi (2000 to 2004), and Florida Department of Environmental Protection's Nutrient Technical Advisory Committee (2003-2006). He presently is an appointed member of the Florida Oceans and Coastal Resources Council, which is charged with developing an annual research plan for the Florida Legislature.

E-mail: khavens@ifas.ufl.edu

Phone: 352-392-9617 ext 232

Curriculum Vitae - includes all publications

Dr. Havens has a website on Zooplankton Ecology: http://planktonweb.ifas.ufl.edu


Courses Taught:

FAS 6932 (Special Topics) credits, lecture, discussion, and field experiences (no formal laboratory component). This course provides graduate students with a comprehensive overview of classical and contemporary aspects of the biology and ecology of freshwater plankton, with particular consideration of how zooplankton structure and function relates to fisheries and water quality. The course includes both classroom and field experiences, critical evaluation of methods used to sample and analyze zooplankton data, and discussion of important papers from the contemporary peer-reviewed literature. Emphasis is on freshwater plankton, however, selected marine literature is covered and most basic principles apply across the freshwater marine continuum. For more information: (download - MS Word)

Power Point Presentations:

August 2005 SNRE Seminar on Algal Blooms

Phosphorus Wind and Water Levels.pdf

Selected Publications

Havens, K.E., K.R. Jin, N. Iricanin, and R.T. James. 2007. Phosphorus dynamics at multiple time scales in the pelagic zone of a large shallow lake in Florida, USA. Hydrobiologia 581: 25-42.

Johnson, S.G., M.S. Allen and K.E. Havens. 2007. A review of littoral vegetation, fisheries and wildlife responses to hydrologic variation at Lake Okeechobee. Wetlands 27: 110-126.

Engstrom, D.E., S.P. Schottler, P.R. Leavitt and K.E. Havens. 2006. A re-evaluation of the cultural eutrophication of Lake Okeechobee, Florida, using multi-proxy sediment records. Ecological Applications 16: 1194-1206.

Havens, K.E. and T.L. East. 2006. Plankton food web responses to experimental nutrient additions in a subtropical lake. TheScientificWorld Journal 6: 827-833.

Carlson, R.E. and K.E. Havens. 2005. Simple graphical methods for the interpretation of relationships between trophic state variables. Lake and Reservoir Management 21: 107-118.

Havens, K.E., D. Fox, S. Gornak and C. Hanlon. 2005. Aquatic vegetation and largemouth bass population responses to water level variations in Lake Okeechobee, Florida (USA). Hydrobiologia 539: 225-237.

James, R.T. and K.E. Havens. 2005. Outcomes of extreme water levels on water quality of offshore and near-shore regions of a large shallow subtropical lake. Archiv fur Hydrobiologie 163: 225-239.

Jeppesen, E., M. Sondergaard, J.P. Jensen, K.E. Havens et al. 2005. Lake responses to reduced nutrient loading – an analysis of contemporary data from 35 European and North American long term studies. Freshwater Biology, 50: 1747-1771.

Pinto-Coelho, R., B. Pinel-Alloul, G. Methot and K.E. Havens. 2005. Crustacean zooplankton in lakes and reservoirs of temperate and tropical regions: variation with trophic status. Canadian Journal of Fisheries and Aquatic Sciences 62: 348-361.

Work, K.A., K.E. Havens, B. Sharfstein and T.L. East. 2005. How important is bacterial carbon to planktonic grazers in a turbid subtropical lake? Journal of Plankton Research 27: 357-372.


The role of freshwater zooplankton (microscopic animals) in lake food webs is a focal area of Dr. Havens' research. He has examined how zooplankton feed on different types of algae, how they respond to various chemical stressors, and how they are affected by predators, including fish. These photo-micrographs show two different species of zooplankton commonly found in Florida lakes ­ Diaptomus dorsalis, a copepod (on the left), and Eubosmina coregoni, a cladoceran (on the right). These animals feed on bacteria, algae, and protozoa. Each animal is less than a millimeter long, yet a liter of lake water may contain hundreds of individuals. Some fish depend on zooplankton as a food source during early stages of their life cycle, so information about the dynamics of zooplankton in a lake can be critical to understanding the population dynamics of fish.

Phytoplankton (suspended algae) experience blooms in Florida lakes, rivers, and coastal waters when there are high concentrations of nutrients and adequate light for net algal growth. Blue-green algae (cyanobacteria) often are the dominant species during blooms in fresh water. Drs. Havens and Phlips have conducted research to examine how dissolved nutrients and light influence biomass and taxonomic composition of phytoplankton, as well as the factors that cause them to bloom in shallow Florida lakes. This photograph shows a cyanobacteria bloom in Lake Okeechobee.

Research conducted by Dr. Havens and his colleagues has shown that submerged plants play a critical role in shallow Florida lakes, providing habitat for fish and other wildlife, and also maintaining clear water by a variety of mechanisms. The plants provide a substrate for periphyton that removes nutrients from the water, thereby indirectly reducing the biomass of phytoplankton. Plants also stabilize sediments, reduce shearing stress on the lake bed from wind-driven waves, and when they carry out intense photosynthesis, result in co-precipitation of phosphorus with calcium at high pH. This photograph shows clear water over a dense bed of submerged plants.

Sandy substrate and a diverse aquatic plant community provide good conditions for foraging and nesting by fish such as largemouth bass. Dr. Havens and colleagues at the Florida Fish and Wildlife Conservation Commission have studied how variations in water level affect aquatic plants and largemouth bass recruitment in Lake Okeechobee. This photograph shows a fish bed in a shallow shoreline area of that lake.


In his current research program, Dr. Havens is working with colleagues in China, Denmark, Italy and Japan to investigate relationships between trophic state parameters (chlorophyll, phosphorus, nitrogen and transparency) in large shallow lakes of the world, and how those relationships are affected by zooplankton grazing pressure, abiotic turbidity and external forcing functions including strong wind events. Lakes in this study include: Okeechobee and Apopka (Florida), Trasimeno (Italy), Taihu and Donghu (China), Kasumigaura (Japan), and Arreso (Denmark).

The two photographs to the right were taken during summer 2006 at Lago Trasimeno, a shallow 125 square kilometer lake in Umbria, Italy. This lake is home to the Italian Institute of Hydrobiology (University of Perugia), where aquatic scientists including Baldi, Vollenweider, and Margalef conducted classical studies of lakes in the early to mid 1900s. The Department of Fisheries and Aquatic Sciences recently established a formal cooperative agreement with the University of Perugia to provide opportunities for faculty and students to conduct collaborative research on freshwater ecosystems in Florida and Italy. Contact Dr. Havens for details if you are a student interested in an international opportunity as part of your graduate studies in FAS.

Page revised January 14, 2008

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